Is Toki damaged by captivity?

A few months ago an important paper was published to establish the harm done to the brains of captive elephants and cetaceans. Putative neural consequences of captivity for elephants and cetaceans by Bob Jacobs, et al. provides “a substantive hypothesis about the negative impact of captivity on the brains of large mammals” The paper supplements observations of pathological behavior like chewing on hard surfaces or head-banging against a wall, and biomedical symptoms of stress, and adds neurological damage that likely accompany such symptoms.